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As the Roaring Twenties crumble into the Great Depression, Virginia Kingsley, New York’s toughest and most successful speakeasy owner, is doing just fine; with the whole world falling apart for so many, bootlegging is a flourishing business.

But when a former lover, Rudy, attempts to coerce his way into her business, Virginia sends him packing and her entire family pays the price: Her niece, Laura, is devastated when her infant son vanishes without a trace. Speculation and rumors abound in the media circus that follows the abduction, and the police have few leads, most of which point straight back at Virginia and her fellow bootleggers.

Suspicious of Rudy but unable to go to the police, Virginia pursues her own investigation, shaking down, threatening, and killing one petty crook after another as she races against time to find her missing great-nephew. In the mean time, Laura's lover and Herald Tribune reporter, Erich, tries to convince Laura that Virginia knows more than she is telling... and may be behind the boy's disappearance.

With time flying past and hope thinning, Virginia, Laura, and Erich must set aside their differences and work together in one last, desperate bid to find the missing child and whoever really abducted him.